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Ellie (Queen Latifah) and Manny (Ray Romano) are expecting their first child, and Manny is obsessed with making life perfect and safe for the family, since his first experiences as a husband and father went bad when his family was killed by hunters. At the same time, Diego (Denis Leary) finds himself unable to catch a cocky gazelle (Bill Hader) he has been stalking and decides to leave the herd, believing that he is losing his predatory nature as a tiger. Sid (John Leguizamo) grows jealous of Manny andEllie and "adopts" three apparently abandoned eggs that he finds in an icy underground cavern and call them Eggbert, Shelly, and Yoko. Manny tells him to put them back, but Sid instead looks after the eggs, which hatch into baby Tyrannosaurus the next morning.

Although Sid tries his best to raise the three dinosaurs, their rambunctious behavior scares away all the other animals' young and ruins a playground Manny built forEllie's baby. A female Tyrannosaurus, Momma, whose eggs Sid stole, soon returns and carries both Sid and her young underground, with Diego in pursuit. Manny, Ellie , Crash, and Eddie (Seann William Scott, Josh Peck) follow as well and discover that the icy cavern leads to a vast jungle populated by dinosaurs thought to be extinct. Here, an Ankylosaurus threatens the herd despite Diego's efforts to fend it off; they are saved from a further crowd of angry reptiles by an insane, one-eyed weasel named Buckminster, or Buck (Simon Pegg).

Buck has been living in this jungle for some time and is chasing Rudy, a large albino Baryonyx, with the intention of avenging the loss of his left eye at Rudy's hands. He agrees to lead the herd through the jungle's perils to Lava Falls, where Momma has taken Sid and her babies. At one point, they have to cross the "Chasm of Death" which if filled with gas fumes (a mixture of helium and laughing gas, causing anyone who breathes in it to laugh uncontrollably while speaking in a high-pitched voice). In the meantime, Sid and Momma try to outdo each other in feeding the offspring; he loses this contest, but is soon welcomed into the family regardless. The next day, however, Sid is separated from the family and attacked by Rudy. Sid is knocked onto a loose rock slab that is floating on a river of lava and about to plummet over the falls.

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